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Punctuated Equilibrium and Memetic Accumulation
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A few posts ago I talked about memetic evolution. As a result of this post one of my readers, Mark, and I had an in depth discussion about what mechanism, exactly, I was trying to describe and whether there really is such a thing as memetic evolution. Mark is a scientist specializing in oncology research (he also has a blog, which you should check out) and he pointed out that evolution is exceptionally complicated and that many people use the term to describe lots of things that aren’t actually evolution by natural selection. Particularly when they’re trying to use it by way of analogy which I was. As part of our discussion a lot of things were clarified for me, and I think I’ve tightened up the analogy and hopefully gotten rid of most of the issues Mark pointed out. This post is about sharing the additional insights which came out of that discussion.
Mark was, of course, correct, there are in fact lots of pitfalls involved in the discussion of evolution and selection, and even if you manage to avoid making any big mistakes there are still numerous specifics that can trip you up as well. For example, most people don’t realize that there are two competing theories regarding the rate at which evolution occurs. And the difference between these two theories turns out to be very important. Not only in general but also for the point I want to make.
The first theory, and the one initially put forward by Darwin, is phyletic gradualism. Under this theory the creation of new species happens very gradually, almost imperceptibly as small changes accumulate over tens of thousands of years. Because of how gradual this process is, you might not end up with a clear line where you can say that one species has changed into another, and, insofar as a layman thinks about evolution with any rigor, they probably envision it working something like this.
The second theory, which was proposed only in 1972, by Niles Eldredge and Stephen Jay Gould, is called punctuated equilibrium. This theory holds that species appear fairly suddenly in response to some rare and geologically rapid event (the punctuation) and that once a species appears that it ends up being relatively stable (the equilibrium). To reiterate, I’m no expert, but it’s my impression that this theory has the most support among scientists, particularly when you’re talking about the big evolutionary events, like speciation. To be clear both kinds of evolution, gradual and punctuated, appear to be taking place, but the latter is more impactful, and more important, particularly when the survival of a given species is really in question.
Having, hopefully, grounded our understanding and discussion of evolution on a somewhat firmer footing, we are still left with the question of how much of that understanding and discussion maps cleanly to the topic of cultural evolution, and beyond that to the more speculative topic of memetic evolution. For instance, insofar as cultural evolution is adaptive, is this adaptation gradual? Or does it operate more along the lines of the punctuated equilibrium model? I’m not entirely sure what would count as hard data when considering these questions, but at the level of anecdote, I’m inclined to believe that the situation is similar to genetic evolution, both forms occur, but that the cultural selection which occurs gradually ends up being less impactful than cultural selection which happens at times of rapid change and extreme crisis.
As I said this is mostly at the level of anecdote, but consider the example of Germany. It’s hard to argue that Germany didn’t have a long martial tradition, starting with their first appearance in the records of the Roman Empire and continuing down through the centuries to the two World Wars. Would you say they still have that culture today? I think most people would agree that they don’t, and that it all changed during the extreme crisis at the end of World War II. Sure there have been many gradual changes to German culture over the years, but the fact that there’s also numerous long-standing stereotypes about Germans would seem to indicate that a relatively stable equilibrium existed as well. From where I sit, this example has all the elements you’d expect if cultural evolution also happened according to the punctuated equilibrium model.
Another example would be the creation of the United States of America. Evolution through natural selection concerns itself with the creation of new species. The parallel in cultural evolution would be the creation of a new culture or nation, and this is an example of exactly that. And, once again, it happened over the course of a few years where things were rapidly changing under crisis conditions. Additionally what resulted was not some incremental change in English culture (though there are obvious connections) but an entirely new culture forged in the fires of the Revolutionary War and the many debates over governments and rights.
The more I consider the question the more I am convinced that there are numerous examples of punctuated equilibrium with respect to cultural evolution. I suspect all of the examples of nations in crisis given by Jared Diamond in his recent book Upheaval (see my review), would end up being examples of the punctuated equilibrium model of cultural evolution as well. And of course these are successful “mutations”, if cultural evolution is anything like biological evolution most mutations are going to end in failure. Is that perhaps the best way of describing communism and fascism?
Obviously not all cultural changes are so large, as I said, I’m sure that things also change gradually, but we would appear to have less to fear from those changes. Sure the vast majority will fail just like all “mutations”, but that failure should be much easier to recover from. Much less disastrous than the analogous “speciation” of adopting something like communism.
If you’ve followed me this far and you accept (even if only for the sake of argument) that punctuated equilibrium applies not only to biological evolution, but to cultural evolution as well, then we’re finally ready to revisit memetic evolution, though right off the bat I’m going to dump the word “evolution”. One of Mark’s bigger contributions in the discussion we ended up having was to point out that once we’ve reached this point that things have been stretched so far that using the term evolution conceals more than it reveals, particularly if we’re more interested in the punctuated equilibrium model of evolution. So we need a new term, but before we get to that what exactly are we talking about? And, in what sense are we talking about something separate and interesting?
When considering the punctuated equilibrium model most of the attention ends up on the punctuation part, but what’s happening during the equilibrium part? Here I’m going to quote liberally from Mark:
[The punctuated equilibrium model] posits [that] major selection events might be somewhat uncommon. As such, we would expect to see accumulations of multiple different mutations, all present in a species' gene pool simultaneously. The longer the period of time free of selection, the greater the potential for diverse new mutations within the species. Since anything directly lethal is going to weed itself out fairly quickly, this enriches for potentially-beneficial mutations. With all these mutations lying around, it's possible for individuals to even have two or more traits that might not be adaptive on their own but that function very well together. This period of stability can be thought of as 'good' in that is allows for much greater variability to enter the population.
Along comes the selection event - the filter, removing anything that can't pass through a particular challenge - and most of that diversity disappears. However, since the population experienced a long period of growth and mutation without being subject to a filter, it's possible that the adaptation that made it through the filter is more complex - is a bigger change - than the kind of single-mutation adaptation you would see from a series of rapid filters. Populations that instead pass through serial filtering events will only be able to select based on single-mutation traits.
….We expect to have multiple possible pro-adaptive traits at any given time, waiting to pass through the next, unexpected, filter and join future generations. Thus, memetic evolution is simply a sub-process of cultural evolution. It would be as meaningless to speak of it in isolation as it would be to talk about accumulating mutations prior to selection events (filters) when speaking of biological evolution.
...Memetic 'evolution' is simply another name for cultural evolution prior to selecting events.
Some of this is obviously speculative, but on the whole Mark’s comments were fantastic, and really helped me to understand something that had previously eluded me, and I agree with everything he said, with one exception… I don’t think it’s “meaningless to speak of it in isolation”. I think “it” is very important to talk about. What is “it”? What is this thing that’s worth discussing, but which is not evolution? I’m going to call it “memetic accumulation”.
For most of history the rate of accumulation for genetic mutations has probably been fairly static. I assume that during periods of greater radiation (if any) that it might have increased, or perhaps the greater the variety of life the greater the space for mutations to occur and perhaps there are other factors as well, but I don’t see any evidence that there were periods where it was significantly faster or slower. There is the Cambrian Explosion, but remember we’re talking about the rate of accumulation, not the rate of evolution or of speciation, and while it was an “explosion” for many things, I don’t think it was an explosion in the accumulation of mutations. In other words I think the rate of mutation accumulation with natural evolution has been pretty constant.
Even when humans entered the scene and started the selective breeding of domesticated animals, this didn’t change the mutation rate, even for the animals in question. (CRISPR, however may be another matter.) We just introduced a lot more filters and selection events. So, if mutations are relatively constant in natural evolution, what about cultural evolution? Has that rate also been constant? I would argue that it hasn’t, and this, more than anything else, is why it’s worth discussing. I suppose, given the fact that humans can introduce new ideas, new potential memes into the space of culture whenever they feel like it, that there are a great many things which could affect the speed at which memetic accumulation occurs. But certainly technology and progress has to have a large impact on that speed, and almost exclusively in the direction of speeding it up. In fact, “something which speeds up the rate of memetic accumulation” is not a half bad definition of progress. But beyond that, might technology and progress have any other effect than generating ideas quickly?
With the advent of global communication and social media, we are moving ever more rapidly in the direction of creating a single ecosystem for ideas, and I don’t think we’ve fully come to terms with what that means or how it will play out. Certainly ideas propagate faster, and I would also say we end up with a handful of “apex ideas” similar to the idea of an apex predator. Which is to say that we're in a space where a memetically fit idea is able to very quickly outcompete all the other ideas among people susceptible to that idea. (Notice the increase in the number of people who believe in conspiracy theories.) Leading to a stratification at the level of ideas rather than at the level of a community or nation. Basically, social media and global communication have allowed invasive species/ideas to go everywhere.
On top of all this there’s one final thing which needs to be pointed out, humans are more removed from issues of actual survival than ever before. Toss all of this together and we have rapid memetic generation, but which results in a relatively barren collection of a few dominant memes/ideologies, none of which are likely to have anything to do with actual survival. Now I’m aware that this is something of an oversimplification, culture is still complex and varied, and people still worry about survival, but we have nevertheless lost an awful lot of both those qualities.
Finally, if I’ve convinced you that memetic accumulation is speeding up, then even if you disagree with me about everything else, you might at least want to examine what the potential consequences of that are with respect to cultural evolution.
Having examined what the modern state of memetic accumulation is within the equilibrium part of the model, what does all of this mean for the eventual “punctuation”? How does our rapid, barren and superficial method of memetic accumulation play out when we actually run into a selection event? Into rapidly changing crisis conditions? Well that’s hard to say, though none of those elements would appear to be positive.
Just by itself, the rapid part isn’t necessarily bad. Perhaps if culture is moving rapidly, then, by the time the eventual crisis rolls around, we will be in some location uniquely well suited for surviving that crisis, a location we would not have reached had we not been moving so quickly. And certainly if there were a bunch of cultures all speeding off towards their own unique locations we might have some expectation that at least one of these locations would be exactly the spot they should be in, but this is where the lack of variety comes into play, we’re not all choosing different locations where we can survive the potential crisis, we seem to all be journeying as quickly as we can towards a small handful of locations, and the rapid bit means if it’s not the right place we will have gone an awfully long distance in the wrong direction. Furthermore, what do these locations look like? If we were really concerned about survival, they would hopefully be strongholds, but if we don’t factor in survival I would think they’re more likely to end up looking like expensive penthouses. Dwellings which look really nice and are great for entertaining, but also the last location you’d want to be in when the zombie apocalypse starts. There’s obviously still a lot of variety in these dwellings, but can anyone honestly tell me we’re not building a lot more penthouses than strongholds these days?
There also seems to be significant effort being spent on getting people to abandon locations which proved to be strongholds in the past. I think I’ve already said this, but it bears repeating, there are essentially three ways to choose a “location” we can choose them randomly, which is essentially what natural evolution is doing. We can choose one based on whether it sounds good or not, but in this sense, as I already pointed out, we’re probably not choosing a stronghold so much as a nice place to live. Or we can choose one based on what’s worked in the past. Any option where we choose is going to be better than random (one would hope) but it’s not clear to me that “sounds good” is definitely better than “worked in the past” (in fact, I strongly suspect it’s worse) and in any event it’s probably best to have cultures in both types of locations.
To be clear, we don’t know which location will best withstand the eventual crisis, because we don’t know what that crisis will look like, but you could certainly see how changing the way in which memetic accumulation happens could change the likelihood of being in the right location. And I hope we can agree on this, even if you don’t agree with me on exactly how memetic accumulation has changed
But beyond all this, there’s probably more bad news, particularly if you believe Nassim Nicholas Taleb’s contention, that in addition to changing to speed of memetic accumulation, that progress and technology has also changed the nature of potential crises as well. That we have made them less frequent, but in the process we’ve also made them larger. As a real world example, lots of people feel that there is no safe location (both figuratively and literally) if the crisis ends up being full scale nuclear war or runaway climate change (I disagree, but I’ve already covered that in past posts). Both crises that have only been made possible recently.
I will freely admit that I’ve followed a long chain of assumptions to get to this point, but strip all that away and I would contend that the two initial ideas, 1) that cultural evolution also follows a pattern of punctuated equilibrium, and 2) that technology and progress can change the rate at which cultural mutations/memes accumulate, are both pretty solid. And both of those together should be enough to introduce serious uncertainty into any claims that conditions are following some long-term, unstoppable, positive trend.
A couple of final things to think about, which I leave as an exercise for the reader:
Are we at a point of “punctuation” right now? If so how’s it looking?
Could memetic accumulation get so out of whack that it actually causes the crisis?
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